下列哪一项未参与形成跨类囊体膜的质子浓度梯度？

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SQL Error: select id,body,options from ***_enewstiku where sourceid='30528' and id < 2465183 and id<>'2465183' order by id desc limit 5 SQL Error: select id,body,options from ***_enewstiku where sourceid='30528' and id > 2465183 and id<>'2465183' order by id asc limit 5